This project investigate the evolution of diversity of fungi by using the wealth of sequence information accumulated in public sequence databases to reconstruct their phylogeny. The phylogeny will then be used to investigate the processes underlying the diversity of species and traits. Of special interest is the evolution of the root associated mutulaism with plants called ectomycorrhiza (ECM). Many of our stand forming trees such as spruce, pine, birch, and oak are depending on ECM for normal growth. ECM has evolved in many (>60) different fungal groups at different times, but the we still know very little about the details. This project will test different hypothesis about the origin of ECM fungi and the dynamic of their evolution.